Helpful articles and insight into our training.

Denson (2018) argues that fMRI evidence from dog studies does not confirm jealousy, as amygdala activation reflects emotional relevance, not specific feelings.

Rivera & Meyers-Manor (2024) found that dogs did not rescue or react differently to distressed strangers, highlighting the importance of owner presence for empathy.

Dickinson & Feuerbacher (2025) revealed that psychological frustration disrupts search and rescue dogs’ focus and performance more than physical stress.

Dobos & Pongrácz (2024) revealed that working dog breeds learn better from humans using speech cues, unlike independent breeds that rely less on verbal communication.

D’Aniello et al. (2016) showed that dogs follow gestures over spoken words when given conflicting cues, emphasizing gesture-based communication.

Konno et al. (2023) revealed that dogs’ dark eyes likely evolved as a facial adaptation that fosters trust and emotional connection with humans.

Smith et al. (2024) found that African wild dogs possess advanced facial and ear musculature, supporting complex within-pack communication.

Zepeda et al. (2024) observed that small-breed dogs communicate distress through body language and vocal cues, especially whining in males.

Ogura et al. (2020) revealed that dogs gaze more at human hands than at those of other animals, highlighting their adaptation to human cues.

Miatkowska et al. (2025) show that owner education in dog communication fosters empathy, mutual understanding, and better welfare.

Siniscalchi et al. (2018) explored how dogs express themselves using body language, sound, and scent in both human and canine communication.

Ren et al. (2024) identified synchronized brain activity between humans and dogs, showing how neural coupling supports emotional and social connection.

Samet et al. (2022) explored how human–dog bonds are measured and proposed new ways to capture dogs’ emotional investment in the bond.

Lezama-García et al. (2025) analyzed how broken human–dog relationships lead to stray dog crises and proposed welfare-centered preventive strategies.

Roma et al. (2024) found that young Canadians respond uniquely to undesired dog behaviors, shaping emotional wellbeing and attachment.

Menor-Campos et al. (2023) highlighted that supportive human–dog relationships improve emotional wellbeing and quality of life for both species.

Allen & Hogg (2022) showed that close dog–owner bonds mitigate emotional loneliness, particularly family and romantic loneliness, improving wellbeing.

Plata & Montiel (2020) documented how Maya peasant-hunters’ dogs transcend utility, embodying protection, prestige, and spiritual meaning.

Verbeek et al. (2024) reviewed over 700 studies, revealing that while human–dog bonds can enhance wellbeing, they may also cause stress depending on the relationship.

Fallahi et al. (2024) explore 40,000 years of human–dog history, showing how evolving societies transformed dogs’ symbolic, cultural, and economic roles.

Verbeek et al. (2024) revealed that the quality of the human–dog bond affects canine welfare, shaped by both dog and owner characteristics.

Czerwinski et al. (2016) revealed that Australian breeders differ widely in how they select dams, with maternal care and genetic testing often undervalued.

Peťková et al. (2024) found that mixed-breed dogs exhibited higher fear and aggression levels than breeds stereotyped as aggressive, challenging public bias.

Habermaass et al. (2023) identified key microbiome shifts in dogs with chronic liver disease, showing that bile alterations affect intestinal bacteria.

Koyama et al. (2024) demonstrated that a prescription diet containing prebiotics improved gut balance and reduced inflammation in dogs with intestinal disease.

Wilson et al. (2024) examined Bacillus coagulans supplementation in dogs and found it supported stool quality and microbiome stability during diet shifts.

Rhimi et al. (2022) examined how diet influences gut microbes and inflammation in canine IBD, offering insights into therapeutic nutrition approaches.

Alvarenga et al. (2024) showed that extruded corn diets improved digestibility, fecal quality, and gut health markers in adult dogs compared to other processing methods.

Riemer (2023) reviewed evidence-based methods for treating and preventing canine noise fears, emphasizing counterconditioning and medication efficacy.

Grigg et al. (2021) revealed that many dogs fear everyday noises at home and that owners frequently misinterpret these stress behaviors as amusement.

Raghy et al. (2023) examined how noise affects dogs’ physiology and behavior, revealing that one-third of dogs suffer from noise aversion.

Sarviaho et al. (2019) discovered genetic regions linked to fear of noises and strangers in dogs that mirror human anxiety-related loci.

Salonen et al. (2020) identified high rates of anxiety among Finnish dogs, highlighting noise sensitivity, fear, and genetic influences across breeds.

Bremhorst et al. (2024) examined the causes, symptoms, and treatments of noise-related fear in dogs, emphasizing early intervention and owner education.

Heurlin et al. (2024) revealed that dogs living closely with humans synchronize their movements with owners, unlike wolves or pack-living dogs.

Dawson et al. (2025) explored how Australian dog breeders choose breeding dogs, rear litters, and socialize puppies, shaping their behavior and future welfare.

Dendoncker et al. (2019) revealed major differences between breeder types, showing that small-scale breeders offer better enrichment and socialization for puppies.

McEvoy et al. (2022) reviewed current knowledge on dog socialization, finding outdated evidence and calling for modern research on optimal timing and breed differences.

Pirrone (2020) highlights the need to educate prospective dog owners on proper puppy acquisition and early socialization to reduce behavior-related relinquishment.

Stolzlechner et al. (2022) showed that structured early challenges make puppies bolder and more resilient, though effects diminish by six months without follow-up.

Abdai et al. (2022) reveal that while dogs retain memories of past interactions, they do not recognize specific humans or robots after brief encounters.

Vervaecke et al. (2021) showed that detection dogs could partially generalize the scent of wolf scat from limited training, supporting their use in ecological monitoring.

Cook (2017) proposes a neuroscience-based approach to studying dog emotion, emphasizing physiology and cognition over assumptions of human-like emotional states.

Corridan et al. (2024) suggest that trauma-informed care, adapted from human psychology, could transform how veterinarians and caregivers assess and treat anxious or reactive dogs.

Fattah and Abdel-Hamid (2020) demonstrated that reward-based methods improved trainability and welfare in German Shepherd police dogs, while punishment caused stress and behavioral issues.

Fonseca et al. (2023) showed that a trainer’s speech tone affects dogs’ and wolves’ emotional and behavioral responses during training, with friendly voices promoting cooperation.

Roz Pooley (2024) described a case of a male Golden Retriever showing aggression toward his owner, which ceased once pain was managed through targeted analgesia.

Ferreira et al. (2022) analyzed stress responses in dogs during grooming and discovered behavioral and physiological changes peaked at arrival and drying but homeostasis was preserved.

Puglisi et al. (2022) tested a novel gel formulation of dog appeasing pheromone (DAP) and observed behavioral calming effects in dogs while waiting for veterinary exams, though physiological stress markers remained unchanged.

Bhadury and Bhattacharjee (2024) synthesized research on how dogs communicate emotions and stress through postures, tail movement, vocal cues, and facial expressions, emphasizing welfare and understanding.

Faragó et al. (2017) found that humans can distinguish emotional meanings in dog growls, which vary across play, threat, and food-guarding contexts—indicating flexible communication strategies.

Scanlon et al. (2020) explored how homeless people in the UK form deep, reciprocal bonds with their dogs—relationships that provide emotional support yet restrict access to housing and aid.

Merkouri et al. (2022) found that while close dog–owner bonds enhance feelings of support and companionship, they may also relate to higher anxiety and depression in owners.

Brucks et al. (2017) found that dogs only show frustration over unequal rewards when an experimenter is present, implying human interaction is crucial to trigger inequity awareness.

Wallis et al. (2020) revealed that dogs with higher obedience levels showed reduced aggression and fearfulness and increased responsiveness, highlighting training’s key role in shaping canine personality.

Cimarelli et al. (2020) found that dogs trained with only partial rewards showed slower progress and developed a pessimistic bias, suggesting welfare concerns in inconsistent clicker training.

Sechi et al. (2016) demonstrated that a nutraceutical diet improved neuroendocrine balance and reduced oxidative stress in dogs suffering from anxiety and chronic stress.

Woszczyło et al. (2020) found that urine scent in estrous female dogs serves as a close-distance cue for males, involving both volatile and nonvolatile compounds rather than long-range pheromones.

Kasuga & Ikeda (2020) found that while owners preferred humanoid robots, dogs showed more social behavior toward a dog-shaped smart speaker, highlighting divergent human–dog perceptions of technology.

Jackson et al. (2021) demonstrated that dogs use olfactory cues to identify larger food quantities, showing no effect of distance or ratio on their scent-based decisions.

Konno et al. (2016) found that ancient dog breeds, genetically closer to wolves, are less likely to make and maintain eye contact with humans compared to modern working and companion breeds.

Lenkei et al. (2019) revealed that Czechoslovakian Wolfdog puppies were more active and exploratory than Labradors, and rearing conditions affected how puppies engaged with humans.

Pongrácz et al. (2004) found that dogs rely on continuous verbal attention during demonstrations, proving communication is key for social learning between dogs and humans.

Fattah and Abdel-Hamid (2020) showed that reward-based training leads to greater olfactory performance and welfare in police dogs, while punishment-based methods harm behavior and wellbeing.

Chen (2017) proposed that dogs and their human companions exhibit hormonal synchronization, with stronger owner–dog bonds linked to greater cortisol alignment.

Foraita et al. (2023) validated the Dog Executive Function Scale across young, adult, and senior dogs, revealing age- and training-related changes in self-control and memory.

Russo et al. (2021) surveyed 111 U.S. animal welfare organizations and found that community programs addressing medical and behavioral needs help retain dogs in homes.

Utami et al. (2019) evaluated Bali’s community-based “Program Dharma,” which boosted rabies vaccination and improved dog welfare in the Sanur sub-district.

Gunter et al. (2019) found that one- and two-night fostering programs lowered shelter dogs’ cortisol levels, showing short-term fostering offers measurable stress relief.

Biswas et al. (2024) found that urban free-ranging dogs in India adjust behavior at seasonal fairs, balancing risk aversion and resource access through cognitive adaptation.

Riemer et al. (2020) reviewed strategies to make veterinary visits less stressful for dogs and cats, emphasizing desensitization, counterconditioning, and gentle handling.

Miller et al. (2019) found that underweight dogs were not more likely to show food aggression toward humans, refuting the belief that past starvation causes such behavior.

Gobbo and Šemrov (2021) analyzed 400 dog-biting incidents, revealing that most bites happened during unprovoked, fast movements near dogs rather than playful contact.

Poncet et al. (2005) discovered that many brachycephalic dogs with upper airway disease also have gastrointestinal tract lesions, revealing a respiratory–digestive connection.

Lindåse et al. (2021) reported that nearly one-third of Swedish show dogs were overweight, but heavier dogs did not perform better in competitions.

Varney et al. (2019) showed that younger, male, and leaner Labradors have higher metabolic rates, while temperature extremes also increase energy demands.

Gobbo and Šemrov (2022) found that aggressive dogs, particularly police dogs, displayed lower self-control in reward delay tasks, linking impulsivity to aggression.

Pan et al. (2018) demonstrated that a diet enriched with medium-chain triglycerides and brain-protective nutrients improved clinical signs of canine cognitive dysfunction.

Canejo-Teixeira et al. (2020) analyzed 255 owner–dog pairs and developed predictive models identifying human and canine traits associated with dysfunctional relationships.

Peter Singer (2018) discusses findings by Cook et al. showing amygdala activation in dogs watching their caregivers feed another dog, suggesting emotional continuity with humans.

Wilson, Soulsbury, and Mills (2024) reveal inconsistencies among clinical animal behaviorists in differentiating fear and frustration, urging a unified framework for emotion-based behavioral assessment.

Martvel and Riemer (2025) developed the first automated AI system to analyze dog facial expressions in real-world settings, finding ear and mouth movements key to emotion detection.

Johnson and Wynne (2022) analyzed 100 U.S. dog trainers’ websites, revealing that language use, gender, and certification differ sharply between aversive and reward-based training methods.

Feuerbacher and Wynne (2016) showed that dogs find access to their owners rewarding, indicating that owner presence can serve as an effective reinforcer in training and relationship-building.

Johnson and Wynne (2024) compared e-collar and food-reward training to stop chasing behavior, finding e-collars more effective under expert control with minimal stress signs observed.

Potter et al. (2021) found that obedience training classes serve as a stealth physical activity intervention, leading owners to walk their dogs more often and take more daily steps while enhancing their bond.

Kuo and Kessler (2024) revealed that dog owners tend to replicate the parenting styles they experienced in childhood, with permissive and protectionistic attitudes influencing caregiving behaviors toward dogs.

Duranton et al. (2017) showed that dogs synchronize their movements, activity, and even gaze direction with their owners, highlighting deep affiliative bonding and interspecies coordination.

Duranton et al. (2019) found that while shelter dogs synchronize walking patterns with their caregivers, they maintain greater distances than pet dogs do with owners, likely due to weaker social bonds.

Grandjean et al. (2022) demonstrated that trained detection dogs can identify SARS-CoV-2 infection from human sweat with up to 94% sensitivity, supporting their role in rapid mass screening.

Salamon et al. (2025) found that while some breeds excelled in scent detection, overall olfactory performance was determined by breed-specific traits rather than group-based selection histories.

Bray et al. (2021) reviewed how refining selection, rearing, and breeding practices can enhance both performance and welfare in working dogs.

Alexander et al. (2016) discovered that soil texture significantly alters the movement of decomposition odor, impacting the accuracy and efficiency of human remains detection dogs.

Turunen et al. (2024) found that dogs trained with progressively weaker scent samples could still detect Eucalyptus hydrolat at astonishingly low concentrations, highlighting the precision of canine olfaction.

Dickinson and Feuerbacher (2025) found that psychological stress from frustration reduces heart rate variability and delays target response times in search and rescue dogs.

Hunter et al. (2020) found major differences between reward-based and balanced trainers in how they view, prevent, and manage canine separation anxiety, including the role of veterinarians.

Foraita et al. (2023) used the Dog Executive Function Scale to show that cognitive skills like working memory and inhibition follow age-related patterns and improve with training.

Castro et al. (2020) showed that dogs trained with aversive-based methods exhibit more stress behaviors, higher cortisol, and pessimistic outlooks compared to reward-trained dogs.

Benítez‐Burraco et al. (2020) suggest that dog domestication and human self-domestication reinforced one another, fostering reduced aggression and the emergence of complex language.

Ren et al. (2024) discovered that mutual gaze and touch trigger synchronized brain activity between humans and dogs, but this neural harmony is disrupted in autism-associated mutant dogs and restored with LSD treatment.

Sexton et al. (2023) revealed that dogs with plainer faces show greater facial expressivity and that humans interpret their emotions more accurately than dogs with complex markings.

Fugazza et al. (2018) found that social learning skills appear in puppies as young as eight weeks old, allowing them to remember actions performed by humans and conspecifics for up to an hour.

Range and Virányi (2013) found that both wolves and dogs can learn from human and canine demonstrators, but domestication has subtly shaped how dogs interpret social cues.

Siniscalchi et al. (2018) reviewed how dogs convey meaning through body posture, scent, sound, and touch, showing that some signals evolve new meanings in communication with humans.

Tonoike et al. (2021) identified genes involved in glucocorticoid function that may underlie the evolution of dogs’ social communication skills with humans during domestication.

Bognár et al. (2023) discovered a general cognitive factor in dogs encompassing learning and problem-solving domains, linked to age, training, personality, and body condition.

Junttila et al. (2022) found that different dog breeds show distinct strengths in social cognition, self-control, and problem-solving—traits likely shaped by selective breeding for diverse working roles.

Lazarowski et al. (2020) tracked the cognitive development of candidate detection dogs, finding steady improvement in inhibitory control, attention, and spatial cognition through early growth.

Holland (2021) explored relationships between medical detection dog trainers and their dogs, showing how emotional ambivalence and mutual influence shape interspecies knowledge during training.

Jones (2023) demonstrated that cooperative care—where dogs can consent to or pause procedures—reduces fear and builds trust during vaccinations, improving both welfare and safety.

Gugołek et al. (2014) compared wild and farmed foxes and raccoon dogs, finding that domestication and controlled feeding increased nutrient digestibility and nitrogen retention in farmed canids.

Jiménez (2021) found that advanced glycation end-products (AGEs) in the blood remain stable regardless of age or size, suggesting conserved metabolic regulation between wild and domestic canids.

Griffin et al. (2023) created a structured hierarchy of dogs’ needs, modeled on Maslow’s framework, to help humans better recognize and fulfill dogs’ true welfare requirements.

Kasprzak and Bornemark (2024) analyzed search-and-rescue dog–handler cooperation through the lens of semiotics, describing teamwork as an inter-species “melody” built on trust, perception, and mutual adaptation.

Heys et al. (2023) found that canine enrichment feeding—using toys, chews, or puzzle feeders—was linked to improved behavior, reduced begging, and higher perceived quality of life in dogs.

Corridan et al. (2024) explored how trauma-informed care principles from human psychology could improve understanding, diagnosis, and treatment of anxiety and fear-based disorders in dogs.

Dare and Strasser (2023) found that calming enrichment, especially lavender scent and tactile items, effectively reduced stress-related behaviors in kenneled shelter dogs during morning cleaning.

Thumpkin et al. (2024) analyzed adopters’ long-term experiences with newly adopted dogs, revealing that patience, trust-building, and safety awareness are essential to successful transitions.

Gemino (2025) analyzed the traits and competencies of Philippine Coast Guard K9 handlers, revealing that conscientiousness, emotional readiness, and trust in dogs are critical for success.

Scalia et al. (2017) found that low-stress handling techniques during veterinary visits lowered fear-related behaviors in dogs, promoting calmer, more positive interactions with veterinarians.

Flint et al. (2024) established key behavioral and physiological markers for assessing positive and negative emotional states in dogs, advancing evidence-based welfare science.

Bradley et al. (2021) discovered that Norfolk Terrier puppies need far less dietary energy for healthy growth than the NRC (2006) equation suggests, highlighting the risk of overfeeding small breeds.

Cocco et al. (2023) found that variations in serotonin, dopamine, and thyroid hormone levels correspond to behavioral differences across dog breeds, offering insights into temperament and selection.

Reynolds et al. (1997) found that sled dogs given carbohydrate supplements immediately after exercise recovered muscle glycogen nearly twice as fast as unsupplemented dogs.

Porter et al. (2021) highlight the need for healthcare providers to assess veterans’ needs, symptoms, and functioning before recommending a service dog or emotional support animal for PTSD.

Riemer (2023) reviewed treatments for canine noise fears and found strong evidence supporting preventive training and counterconditioning, while most alternative remedies remain ineffective.

Flint et al. (2023) discovered that long-lasting chews were more effective than treat-dispensing toys in calming dogs left briefly alone, promoting emotional wellbeing.

Vasconcellos et al. (2016) found that training sessions with human caretakers lowered cortisol levels in wolves and dogs alike, showing shared benefits of structured interaction.

Wyrick et al. (2025) found that gray wolves in less-crowded areas investigate simulated intrusions more intensely, showing adaptive territorial strategies.

Root-Gutteridge et al. (2021) discovered that dogs react faster and more intensely to puppy-like distress cries than to human infant cries, revealing limits of cross-species empathy.

Bhattacharjee et al. (2017) found that free-ranging dogs show developmental changes in understanding human pointing cues, suggesting experience-based learning.

Cafazzo et al. (2018) show wolves reconcile post-conflict to sustain cooperation, whereas domestic dogs avoid contact with opponents.

Range & Virányi (2014) found that wolves surpass dogs in imitating conspecifics, supporting the view that wolf cooperation influenced domestication.

Marshall-Pescini et al. (2018) reveal that wolves cooperate more effectively than dogs in problem-solving tasks, reflecting their differing social ecologies.

Range et al. (2019) show that wolves lead and dogs follow in cooperative tasks with humans, supporting the idea that domestication favored deference.

Werhahn et al. (2016) reveal that wolves and dogs both follow conspecifics’ gaze, but differ in how they respond to human gaze, reflecting domestication effects.

Range & Virányi (2013) found that dogs and wolves benefit from social learning with human or conspecific demonstrators, highlighting nuanced effects of domestication.

Lampe et al. (2017) reveal that wolves excel over dogs in causal cognition tasks, suggesting domestication has reshaped specific problem-solving skills.

Mantar & Demirel (2024) review recent findings on herbal supplements used to support dogs with cognitive dysfunction, highlighting potential symptom relief.

Nokay (2023) highlights the potential of combining traditional Chinese food therapy with modern dietary strategies to manage canine cognitive dysfunction.

Haake et al. (2024) compared standard CCD questionnaires, showing CADES detects early cognitive decline more effectively than CCAS or CCDR.

Kim et al. (2024) developed machine learning models using blood biomarkers to detect canine cognitive dysfunction early, offering new diagnostic potential.

Merbl et al. (2025) investigated blood–brain barrier dysfunction in dogs with CCD using MRI subtraction enhancement, finding mixed results.

Noche et al. (2024) found that behavioral enrichment boosted hippocampal volume and helped counter brain atrophy in aging beagles, paralleling Alzheimer’s research.

Dewey et al. (2020) report that dogs with cognitive dysfunction show hippocampal atrophy on MRI, paralleling Alzheimer’s pathology in humans.

Mihevc et al. (2020) found that nitrosative stress and nitric oxide synthases may drive neurodegeneration in dogs with cognitive dysfunction.

Osburn et al. (2024) report that canine cognitive dysfunction involves Alzheimer’s-like brain transcriptome and extracellular vesicle RNA changes.